England, early 20th century. The future writer and philologist John Ronald Reuel Tolkien (1892-1973) and three of his schoolmates create a strong bond between them as they share the same passion for literature and art, a true fellowship that strengthens as they grow up, but the outbreak of World War I threatens to shatter it.
When Gerda Wegener asks her husband Einar to fill in as a portrait model, Einar discovers the person she's meant to be and begins living her life as Lili Elbe. Having realized her true self and with Gerda's love and support, Lili embarks on a groundbreaking journey as a transgender pioneer.